/* 生物形态设计 - 炭黑主题 */
:root {
    --charcoal: #1a1e24;
    --deep-moss: #2a3f36;
    --amber: #ff9e3d;
    --lichen: #b5bf50;
    --mist: #e0e6eb;
    --organic-curve: 35px;
    --fluid-transition: all 0.5s cubic-bezier(0.65, 0, 0.35, 1);
}

* {
    margin: 0;
    padding: 0;
    box-sizing: border-box;
    font-family: 'Segoe UI', Roboto, Oxygen, Ubuntu, Cantarell, 'Open Sans', sans-serif;
}

body {
    background-color: var(--charcoal);
    color: var(--mist);
    line-height: 1.7;
    background-image: 
        radial-gradient(circle at 10% 20%, rgba(42, 63, 54, 0.2) 0%, transparent 20%),
        radial-gradient(circle at 90% 80%, rgba(181, 191, 80, 0.1) 0%, transparent 25%);
}

a {
    text-decoration: none;
    color: var(--lichen);
    transition: var(--fluid-transition);
}

a:hover {
    color: var(--amber);
    text-shadow: 0 0 8px rgba(255, 158, 61, 0.4);
}

.container {
    width: 92%;
    max-width: 1300px;
    margin: 0 auto;
}

/* 仿生头部设计 */
header {
    background-color: rgba(26, 30, 36, 0.92);
    padding: 25px 0;
    position: sticky;
    top: 0;
    z-index: 1000;
    backdrop-filter: blur(8px);
    border-bottom: 1px solid rgba(42, 63, 54, 0.3);
    border-radius: 0 0 var(--organic-curve) var(--organic-curve);
}

.header-content {
    display: flex;
    justify-content: space-between;
    align-items: center;
    flex-wrap: wrap;
}

.logo {
    font-size: 28px;
    font-weight: 700;
    background: linear-gradient(135deg, var(--lichen), var(--amber));
    -webkit-background-clip: text;
    -webkit-text-fill-color: transparent;
    letter-spacing: 1.5px;
}

nav ul {
    display: flex;
    list-style: none;
    gap: 25px;
}

nav ul li a {
    padding: 10px 20px;
    border-radius: 50px;
    background: rgba(42, 63, 54, 0.3);
    font-weight: 500;
    display: inline-block;
    transform: scale(1);
}

nav ul li a:hover {
    background: var(--deep-moss);
    transform: scale(1.05);
    box-shadow: 0 4px 12px rgba(42, 63, 54, 0.4);
}

/* 流体内容区域 */
.main-content {
    background: rgba(26, 30, 36, 0.85);
    border-radius: var(--organic-curve);
    padding: 40px;
    margin: 40px 0;
    backdrop-filter: blur(6px);
    border: 1px solid rgba(42, 63, 54, 0.3);
    box-shadow: 
        0 6px 18px rgba(0, 0, 0, 0.3),
        inset 0 1px 0 rgba(255, 255, 255, 0.05);
}

.section-title {
    font-size: 26px;
    margin-bottom: 30px;
    padding-bottom: 15px;
    position: relative;
    color: var(--mist);
    font-weight: 600;
}

.section-title::after {
    content: '';
    
    bottom: 0;
    left: 0;
    width: 120px;
    height: 3px;
    background: linear-gradient(90deg, var(--lichen), transparent);
    border-radius: 3px;
}

/* 生物形态卡片 */
.bio-grid {
    display: grid;
    grid-template-columns: repeat(auto-fill, minmax(300px, 1fr));
    gap: 30px;
}

.bio-card {
    background: rgba(42, 63, 54, 0.15);
    border-radius: calc(var(--organic-curve) - 10px);
    overflow: hidden;
    transition: var(--fluid-transition);
    border: 1px solid rgba(42, 63, 54, 0.4);
    position: relative;
}

.bio-card::before {
    content: '';
    
    top: 0;
    left: 0;
    width: 100%;
    height: 100%;
    background: radial-gradient(circle at 70% 30%, rgba(181, 191, 80, 0.1), transparent 60%);
}

.bio-card:hover {
    transform: translateY(-8px);
    box-shadow: 
        0 12px 24px rgba(0, 0, 0, 0.4),
        0 0 0 1px rgba(181, 191, 80, 0.2);
}

.bio-image {
    width: 100%;
    height: 200px;
    object-fit: cover;
    border-bottom: 1px solid rgba(42, 63, 54, 0.4);
}

.bio-content {
    padding: 25px;
}

.bio-title {
    font-size: 20px;
    margin-bottom: 15px;
    font-weight: 600;
    color: var(--mist);
    display: -webkit-box;
    -webkit-line-clamp: 2;
    -webkit-box-orient: vertical;
    overflow: hidden;
}

.bio-meta {
    display: flex;
    justify-content: space-between;
    font-size: 14px;
    color: var(--lichen);
    margin-top: 20px;
}

.bio-meta span {
    display: flex;
    align-items: center;
    gap: 5px;
}

/* 分类标签 */
.bio-tag {
    display: inline-block;
    padding: 5px 15px;
    background-color: var(--deep-moss);
    color: var(--mist);
    border-radius: 50px;
    font-size: 13px;
    margin-bottom: 15px;
    font-weight: 500;
}

/* 文章详情页 */
.bio-article {
    max-width: 850px;
    margin: 0 auto;
}

.article-header {
    margin-bottom: 50px;
    text-align: center;
}

.article-title {
    font-size: 36px;
    margin-bottom: 25px;
    line-height: 1.3;
    color: var(--mist);
    font-weight: 700;
}

.article-meta {
    display: flex;
    justify-content: center;
    gap: 30px;
    color: var(--lichen);
    margin-bottom: 30px;
    flex-wrap: wrap;
}

.article-image {
    width: 100%;
    max-height: 500px;
    object-fit: cover;
    border-radius: var(--organic-curve);
    margin-bottom: 40px;
    box-shadow: 0 8px 24px rgba(0, 0, 0, 0.4);
}

.article-content {
    line-height: 1.9;
    font-size: 18px;
}

.article-content p {
    margin-bottom: 25px;
}

.article-content img {
    max-width: 100%;
    height: auto;
    border-radius: calc(var(--organic-curve) - 10px);
    margin: 30px 0;
    box-shadow: 0 4px 12px rgba(0, 0, 0, 0.3);
}

/* 流体分页 */
.bio-pagination {
    display: flex;
    justify-content: center;
    gap: 20px;
    margin-top: 50px;
}

.bio-pagination a {
    padding: 12px 25px;
    border-radius: 50px;
    background-color: rgba(42, 63, 54, 0.3);
    font-weight: 500;
    transition: var(--fluid-transition);
}

.bio-pagination a:hover {
    background-color: var(--deep-moss);
    transform: translateY(-3px);
}

/* 友情链接 - 仿生群落样式 */
.bio-links {
    background: rgba(42, 63, 54, 0.2);
    border-radius: var(--organic-curve);
    padding: 30px;
    margin: 40px 0;
    backdrop-filter: blur(4px);
    border: 1px solid rgba(42, 63, 54, 0.3);
}

.bio-links h3 {
    margin-bottom: 25px;
    color: var(--mist);
    font-size: 22px;
    position: relative;
    display: inline-block;
}

.bio-links h3::after {
    content: '';
    
    bottom: -5px;
    left: 0;
    width: 100%;
    height: 2px;
    background: linear-gradient(90deg, var(--amber), transparent);
}

.bio-links-container {
    display: flex;
    flex-wrap: wrap;
    gap: 15px;
}

.bio-links-container a {
    padding: 10px 20px;
    background-color: rgba(42, 63, 54, 0.4);
    border-radius: 50px;
    font-size: 15px;
    transition: var(--fluid-transition);
}

.bio-links-container a:hover {
    background-color: var(--deep-moss);
    transform: scale(1.05);
}

/* 页脚 - 菌丝体样式 */
footer {
    background-color: #14181c;
    padding: 40px 0;
    text-align: center;
    margin-top: 60px;
    border-top: 1px solid rgba(42, 63, 54, 0.4);
    position: relative;
    overflow: hidden;
}

footer::before {
    content: '';
    
    top: 0;
    left: 0;
    width: 100%;
    height: 100%;
    background: 
        radial-gradient(circle at 20% 30%, rgba(181, 191, 80, 0.05), transparent 40%),
        radial-gradient(circle at 80% 70%, rgba(255, 158, 61, 0.05), transparent 40%);
}

.copyright {
    font-size: 15px;
    color: var(--lichen);
    position: relative;
}

/* 响应式设计 */
@media (max-width: 992px) {
    .header-content {
        flex-direction: column;
        gap: 25px;
    }
    
    nav ul {
        justify-content: center;
    }
    
    .bio-grid {
        grid-template-columns: repeat(auto-fill, minmax(280px, 1fr));
    }
}

@media (max-width: 768px) {
    .main-content {
        padding: 30px;
    }
    
    .article-title {
        font-size: 28px;
    }
    
    .article-meta {
        gap: 15px;
    }
    
    .bio-pagination {
        flex-direction: column;
        align-items: center;
    }
}